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Generally how does a male animal realize that the female is receptive to copulation?

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In most vertebrate species with internal fecundation the females have reproductive cycles with fertile periods. During this period the female secretes pheromones (odoriferous substances that attract the male of the species) from the skin and mucosae. The presence of the male individual and his pheromones also stimulates the release of pheromones by the female. (Many animals also use pheromones for territorial demarcation and for signal transmission between individuals about the location of dangers and food.)

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