1- Henking- He gave the importance of X body transfer in spermatogenesis of insects
2 – T.H. Morgan – Morgan gave the importance of physical association or linkage of the two genes and coined the term linkage.
3 – Sutton & Boveri – He gave the importance of the pairing and separation of a pair of chromosomes would lead to the segregation of a pair of factors they carried, it is called chromosomal theory of inheritance.