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Species-Area relationship was given by

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Species-Area relationship was given by
A. British zoologist and geographer David Tilman
B. German botanist and geographer Alexander von Humboldt
C. German naturalist and geographer Alexander von Humboldt
D. British naturalist and geographer Alexander von Huboldt
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Species-Area relationship was given by German naturalist and geographer Alexander von Humboldt.
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